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Zacuto's Gorilla Plate V1 Adapter centers the Gorilla Plate V1's offset tripod holes by screwing into the bottom of the plate and providing a set of aligned holes. This is necessary when the Gorilla Plate V1 and Z-Finder are mounted to another manufacturer's baseplate. Without the adapter, the camera won't be centered and the lens will not line up with a matte box or follow focus.
Note: A height adjustable matte box is needed if using a matte box with the Gorilla Plate V1 on another manufacturer's plate system unless, like the Zacuto DSLR Baseplate, their plate system is height adjustable.